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The Skyshrine Guardian is one of the two Avatars you can control in Proving Grounds: The Underdepths.
The Skyshrine Guardian is a berserker, and seems to deal less damage (and thus grant a slower completion time) than the Skyshrine Infiltrator.
Defeating the first boss, Veritor Buldama, awards your third temporary buff ability: Veritor's Deadly Round-up.
Defeating the second boss, Gikrakiis, grants the ability Shed Skin, a self cure.
Defeating the third boss, Aherin the Mighty, grants the ability Aherin's Mighty Slam, a strong attack.
Defeating Spikran Vergun grants you Strength of Skyshrine, a buff which boosts your abilities, and also allows you to cast while moving.
Defeating Kero'Var grants you Kick Up Dust, an Area-Of-Effect attack ability that also increases damage done to the target.
Finally, defeating Galstav, Scorcerer of Night grants you Grounded, a temporary buff that protects you from static storms, which is useful against the final boss, Kildrukaun the Ancient.